Used to decrypt the title key for every Wii U application. Used by the Espresso Boot ROM to decrypt the Cafe OS kernel at load time. See Fail0verflow's talk for more information on the NANDloader. Used by the Espresso Boot ROM to decrypt vWii System Menu and new NANDloader binaries at load time.
After the Boot ROM reads these keys, it disables access until the Espresso is reset again. The Espresso keybank is 64 bytes, with the first 32 being the Ancast Keys.